Lepidoptera Survey



Jing Zhang

Date of this Version


Document Type



Taxonomic Report of the International Lepidoptera Survey (May 1, 2022) 10(4): 1-13

Also available at https://lepsurvey.carolinanature.com/ttr/ttr-10-4.pdf

Also available at the Internet Archive, the Biodiversity Heritage Library, Zogobat, and Zenodo

ZooBank registration: http://zoobank.org/CCF018F4-0188-451A-BE75-006F64938595


Copyright 2022, the International Lepidoptera Survey. Open access material

License: Creative Commons Attribution-ShareAlike-NonCommercial 4.0 International (CC BY-SA-NC 4.0 International)


Two new skipper butterfly (Hesperiidae) species are described from the United States: Staphylus floridus Grishin, sp. n. (type locality in Florida, Volusia County) and Staphylus ecos Grishin, sp. n. (type locality in Texas, Brewster County). They are cryptic and hence escaped recognition. They differ from their sister species by the relative size and morphology of genitalia and by genotype—including and beyond the COI barcode—thus, suggesting genetic isolation that argues for their species-level status. A lectotype is designated for Helias ascalaphus Staudinger, 1876. Staphylus opites (Godman & Salvin, 1896), stat. rest. is a species-level taxon and not a synonym of Staphylus vincula (Plötz, 1886), while Pholisora iguala Williams & Bell, 1940, syn. n. is a junior subjective synonym of S. vincula.

COI = c oxidase subunit I