Natural Resources, School of


Date of this Version



Cederbaum, S.B., B.C. Faircloth, T.M. Terhune, J.J. Thompson, and J.P. Carroll (eds.). 2009. Proceedings of Gamebird 2006. Quail VI and Perdix XII. Warnell School of Forestry and Natural Resources. University of Georgia, Athens. 541pp.


Proceedings of Gamebird 2006: a combined conference of the North American Quail Conference series and the Perdix Series.


Foreword v

Acknowledgements vi

Editorial Panel vii

Sponsors x

Invited Manuscripts 2

Integrating Management, Research, and Monitoring: Balancing the 3-Legged Stool. Michael J. Conroy & James T. Peterson . 2

Management of Southern African Gamebirds: Opportunities and Threats. Tim Crowe . 11

Taking the Northern Bobwhite Conservation Initiative to the Next Level. Donald F. McKenzie . 16

Restoring a Gray Partridge (Perdix perdix) Population and the Future of Predation Control. G.R. Potts . 24

Abundance Estimation 27

The GWCT Partridge Count Scheme: A Volunteer-Based Monitoring and Conservation Promotion Scheme. Julie Ewald, Nevile Kingdon, & Hugues Santin-Janin . 27

Refining the Morning Covey-Call Survey to Estimate Northern Bobwhite Abundance. Joshua P. Rusk, Jason L. Scott, Fidel Hern´andez, & Fred C. Bryant . 38

Estimating Sample Sizes for Distance Sampling of Autumn Northern Bobwhite Calling Coveys. Mark D. Smith, Richard G. Hamrick, L. Wes Burger, Jr., & John P. Carroll 46

Summer Male Call Index Relative to Nesting Chronology and Autumn Density of the Northern Bobwhite. Theron M. Terhune, Richard. G. Hamrick, D. Clay Sisson, & H. Lee Stribling 54

Brood Ecology 66

Invertebrate Biomass and Richness in Various Food Plot Types in East Texas. Charles W. Anderson, R. Montague Whiting, Jr., Donald R. Dietz, & Richard M. Capps 66

Integrating Burning and Insecticide to Reduce Fire Ant Impacts on Bobwhite Chicks. Amy Norton Johnson, C. Brad Dabbert, Robert B. Mitchell, & Harlan G. Thorvilson 78

Northern Bobwhite Brood Habitat Selection in South Florida. Nevena Martin, James A. Martin, & John P. Carroll . . 88

Conservation 99

Managing Black-throated Bobwhite for Sustainability in Belize: Preliminary Results of a Population Study. Jack Eitniear, Reynold Cal, Wilbur Martinez, Omar Fiqueroa, & John Baccus . 99

Delineation of Gambel’s Quail Habitat in the Trans-Pecos, Texas. Alfonso Ortega-Sanchez, Louis A. Harveson, Roel R. Lopez, & Michael R. Sullins . 106

Habitat Use and Survival of the Spotted Tinamou (Nothura maculosa) in Agroecosystems in the Province of Buenos Aires, Argentina. Jeffrey J. Thompson & John P. Carroll 111

Gamebird Genetics 121

Release of Pen-reared Bobwhites: Potential Consequences to the Genetic Integrity of Resident Wild Populations. Kristine O. Evans, Mark D. Smith, Loren W. Burger, Jr., Rachel J. Chambers, Allan E. Houston, & Rick Carlisle 121

Habitat 135

Effects of Sex, Age, and Habitat on Northern Bobwhite Spring Dispersal Patterns. M. Patrick Cook, Richard G. Hamrick, & John P. Carroll 135

Usable Space Versus Food Quantity in Bobwhite Habitat Management. Douglas S. Cram, Ronald E. Masters, Fred S. Guthery, David M. Engle, & Warren G. Montague 146

Winter Food Habits and Preferences of Northern Bobwhites in East Texas. Donald R. Dietz, Jr., R. Montague Whiting, Jr., & Nancy E. Koerth . 160

Habitat Use and Survival of Gray Partridge Pairs in Bavaria, Germany. Wolfgang Kaiser, Ilse Storch, & John P. Carroll 172

Effects of Timber Density on Northern Bobwhite Autumn Abundance. Ian T. Little, Shane D. Wellendorf, William E. Palmer, & John P. Carroll . 178

Usable Space Versus Habitat Quality in Forest Management for Bobwhites. Ronald E. Masters, Fred S. Guthery, W. Russ Walsh, Douglas S. Cram, & Warren G. Montague 184

Influence of Habitat, Fire, and Weather on Bobwhite Abundance at Avon Park Air Force Range, Florida. Brett D. Miley & Marian Lichtler . . 193

Species Visitation at Quail Feeders and Guzzlers in Southern New Mexico. Dale Rollins, Ben D. Taylor, Troy D. Sparks, Tom E. Wadell, & George Richards . . 210

Population Response of Northern Bobwhite to Field Border Management Practices in Mississippi. Mark D. Smith & L. Wes Burger, Jr. . 220

Northern Bobwhite Demographic and Population Response Following an Intensive Habitat Modification to an Agricultural Landscape. Theron M. Terhune, D. Clay Sisson, Steven D. Mitchell, & H. Lee Stribling . 232

Habitat Management 251

Half-Cutting as a Management Tool to Increase Abundance of Northern Bobwhite in South Texas. Dale A. Hall & Nova J. Silvy . 251

Testing the Value of Prickly Pear Cactus as a Nest-Predator Deterrent for Northern Bobwhite. Fidel Hern´andez, Scott E. Henke, Nova J. Silvy, & Dale Rollins 256

The Shepherd Project: A Case Study of Private Management for Ring-necked Pheasants (Phasianus colchicus) in Montana. Bruce Kania & Frank Stewart . 260

Impact of Game Crops Intensification and Hunting Management on Red-Legged Partridge. Franoise Ponce-Boutin, Jean-Franois Mathon, & Tanguy Le Brun 267

Effects of Two Burn Scales on Northern Bobwhite Demographic Parameters and Home Range Size. Shane D. Wellendorf & William E. Palmer . . 271

Effect of Supplemental Feeding onWinter Diet and Body Mass of Northern Bobwhites. Ryan E. Whitelaw, William E. Palmer, Shane D. Wellendorf, & John P. Carroll 282

Policy and Conservation 291

The GWCT Grey Partridge Recovery Programme: a Species Action Plan in Action. Nicholas J. Aebischer . 291

Conservation Reserve Program Grasslands and Ring-Necked Pheasant Abundance in Minnesota. James F. Drake, Richard O. Kimmel, J. David Smith, & Gary Oehlert 302

Methods for ControllingWoody Invasion into CRP Fields in Tennessee. John P. Gruchy, Craig A. Harper, & Matthew J. Gray . 315

Management of Montezuma Quail in Texas: Barriers to Establishing a Hunting Season. Louis A. Harveson 322 Long-Term Climate Trends and Northern Bobwhite Populations in South Texas. Jeffrey J. Lusk, Fred S. Guthery, Markus J. Peterson, & Stephen J. DeMaso 328

Hunting Success on Albany, Georgia Plantations: The Albany Quail Project’s Modern Quail Management Strategy. H. Lee Stribling & D. Clay Sisson . 338

Summary of Georgia’s Bobwhite Quail Initiative 2000-2005. Reggie E. Thackston, Chris D. Baumann, Bobby T. Bond, & Mark D. Whitney 348

Managing Quail in Arizona: Meeting New Challenges with Old Techniques. Mark L. Zornes . . 370

Restocking 380

A Review of Gray Partridge Restocking in the UK and its Implications for the UK Biodiversity Action Plan. Stephen J. Browne, Francis Buner, & Nicholas J. Aebischer 380

How to Re-Introduce Gray Partridges - Conclusions from a Releasing Project in Switzerland. Francis Buner 391

Comparison of Survival, Productivity, Movements, and Habitat Use of Pre-Season Released Quail onWild Northern Bobwhites on Groton Plantation, South Carolina. Derek A. Eggert, Brad S. Mueller, Lamar Robinette, & Shane Wellendorf . 396

Survival and Predation 410

Survival and Habitat Use of Wild Pheasant Broods on Farmland in Lower Austria. Thomas H. Bliss, Brandon C. Anderson, Roger A.H. Draycott, & John P. Carroll . . 410

A Review of Crippling Loss for Northern Bobwhites. Aaron M. Haines, Fidel Hern´andez, Scott E. Henke, & Ralph L. Bingham 420

Fate and Survival of Radio-marked Montezuma Quail. Froylan Hernandez, Eric Garza, Louis A. Harveson, & Clay E. Brewer . . 426

Over-winter Survival of Northern Bobwhite in Relation to Landscape Composition and Structure. R. Douglas Holt, Loren W. Burger, Jr., Bruce D. Leopold, & Dave Godwin 432

An Evaluation of Short-term Mesocarnivore Control for Increasing Hatch Rate in Northern Bobwhites. Eddie K. Lyons, Jason Frost, Dale Rollins, Ben Taylor, & Cody Scott . 447

Survival of Female Scaled Quail During the Breeding Season at Three Sites in the Chihuahuan Desert. Dale Rollins, Ben D. Taylor, Troy D. Sparks, Robert J. Buntyn, Scott E. Lerich, Louis A. Harveson, Tom E. Wadell, & Cody B. Scott 456

Survival and Causes of Mortality for Northern Bobwhites in the Southeastern USA. D. Clay Sisson, Theron M. Terhune, H. Lee Stribling, Jerald F. Sholar, & Steven D. Mitchell . 467

Techniques 480

Necklace-type Transmitter Attachment Method for Ruffed Grouse Chicks. Christopher A. Dobony, Brian W. Smith, John W. Edwards, & Thomas J. Allen 480

A Method for Determining Asymptotes of Home-Range Area Curves. Aaron M. Haines, Fidel Hern´andez, Scott E. Henke, & Ralph L. Bingham 489

Total Body Electrical Conductivity for Determining Carcass Fat in Ruffed Grouse. Aaron B. Proctor & John W. Edwards . . 499 Age Determination of Pheasants (Phasianus Colchicus) using Discriminant Analysis. Maureen I.A. Woodburn, John P. Carroll, Peter A. Robertson, & Andrew N. Hoodless . 505

Telemetry Effects 518 Additional Evidence Against Radio-handicapping of Northern Bobwhites. D. Clay Sisson, Theron M. Terhune, & H. Lee Stribling . 518

Index 526