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In sweetclover (Melilotus alba Desr.) the allelic pair designated B/b was originally thought to be concerned with coumarin biosynthesis. Extracts of BB plants were reported to contain substantial amounts of both free and bound coumarin, while extracts of bb plants contained only bound coumarin (Goplen, Greenshields and Baenzinger 1957; Haskins and Gorz 1957), more recently identified as coumarinic acid glucoside (β-D-glucosyl-cis-o-hydroxycinnamic acid) (Kosuge 1961; Stoker and Bellis 1962). The bound-coumarin character behaved as a simple recessive in crosses between bb and BB plants (Goplen, Greenshields and Baenzinger 1957).