Insecta Mundi is a journal primarily devoted to insect systematics, but articles can be published on any non-marine arthropod taxon. Manuscripts considered for publication include, but are not limited to, descriptions of new taxa, informal taxonomic notes, bibliographies, checklists, catalogs, and reviews. For papers about new insect species, Insecta Mundi provides authors early publication so that the species may be included in other books and catalogs.

This archive of older issues is presented here through the courtesy and cooperation of the Insecta Mundi editorial board. DjVU files were retrieved from the Insecta Mundi Online page at, converted to PDF format, text-recognized with OCR, and split into individual articles, which are posted here with abstracts under their year of publication.

Current issues of Insecta Mundi are available online at
Volumes 17–20 and current issues are available online at

Insecta Mundi is published by the Center for Systematic Entomology, P.O.Box 141874, Gainesville, Florida 32614-1874 USA.


A new species of Biclonuncaria Razowski and Becker, 1993 (Lepidoptera: Tortricidae: Polyorthini) from Costa Rica, John W. Brown

New species and taxonomical and geographical notes on Cerambycidae and Disteniidae (Coleoptera: Chrysomeloidea), Daniel Heffern, Juan Pablo Botero, and Antonio Santos-Silva

Taxonomic notes on the Luciolinae (Coleoptera: Lampyridae), Oliver Keller and Lesley Ballantyne

New host and distribution records plus additional notes for North American species of Chrysobothris Eschscholtz (Coleoptera: Buprestidae), with designation of a lectotype for Chrysobothris vulcanica LeConte, 1861, Richard L. Westcott

Thirteen new species of butterflies (Lepidoptera: Hesperiidae) from Texas, Jing Zhang, Qian Cong, and Nick V. Grishin


Natural history of Cassida sphaerula Boheman, 1854 (Coleoptera: Chrysomelidae: Cassidinae: Cassidini) on Arctotheca prostrata (Salisb.) Britten (Asteraceae: Arctotidinae) in South Africa, with a checklist of South African Cassidinae (leaf-mining and tortoise beetles), Sally Adam, Mariana Campos, Hugh D.C. Heron, C. L. Staines, Rob Westerduijn, and Caroline Simmrita Chaboo

A second species of the genus Erebaces Pascoe, 1871 (Curculionidae: Molytinae: Cryptorhynchini) from the Philippines, Robert S. Anderson

First record of Agrilus cliftoni Knull and Anthaxia viridifrons Gory (Coleoptera: Buprestidae) in Wisconsin, Lawrence Barringer

New species, synonymy, new records, and taxonomic notes in American Cerambycidae (Coleoptera), Juan Pablo Botero and Antonio Santos-Silva

Genomics reveals a new genus and species from a single female specimen (Lepidoptera: Hesperiidae: Hesperiinae: Hesperiini: Moncina), Ernst Brockmann, Jing Zhang, Qian Cong, and Nick V. Grishin

A review of host plants for the tortricid tribe Grapholitini, with a synopsis of host utilization by genus (Lepidoptera: Tortricidae), John W. Brown

New species and new combinations in Afrotropical Eucosmocydia Diakonoff, 1988 (Lepidoptera: Tortricidae: Olethreutinae), John W. Brown, Jozef Razowski, Leif Aarvik, Alicia E. Timm, and Robert S. Copeland

Taxonomic review of the North American dung beetle genus Boreocanthon Halffter, 1958 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini), W. D. Edmonds

A new species of Brachycerocoris Costa (Hemiptera: Heteroptera: Pentatomidae: Podopinae) from the Philippines, Joseph E. Eger Jr.

The multiform genus Psyrassa Pascoe (Coleoptera: Cerambycidae: Elaphidiini): new species, new records, synonyms and transfers, Kimberly García and Antonio Santos-Silva

Descripción de una nueva especie de Tegmaleurodes Martin (Hemiptera: Aleyrodidae) de Guatemala, José Francisco García-Ochaeta

Description of a new species of Aleuroparadoxus Quaintance and Baker (Hemiptera: Aleyrodidae) from Guatemala, José Francisco García Ochaeta and Anil Kumar Dubey

Descripción de dos especies nuevas de Mesoplia Lepeletier (Hymenoptera: Apoidea: Apidae) para Cuba y Las Bahamas, con notas sobre taxonomía y distribución del género en las Antillas, Julio A. Genaro and Dayron Breto

The first report of Lissomus Dalman (Coleoptera: Elateridae: Lissominae) from the Greater Antilles, with two new species from the Dominican Republic, Paul J. Johnson

On the firefly (Coleoptera: Lampyridae) species of Carl Linnaeus, Oliver Keller

Identification of a potential pathway of the exotic black weevil (Coleoptera: Curculionidae) in South Korea, Sohee Kim, Seon Woo Lee, and Soo-Jung Suh

Geographic variation in host selection in the spider wasps Entypus unifasciatus (Say) and Tachypompilus ferrugineus (Say) (Hymenoptera: Pompilidae), II, Frank E. Kurczewski, Joseph W. Stoll, Rick C. West, Kelly C. Kissane, Paige R. Chesshire, and Neil S. Cobb

Host selection and nesting behavior of Nearctic trapdoor spider-hunting spider wasps (Hymenoptera: Pompilidae: Pepsinae, Pompilinae), Frank E. Kurczewski and Rick C. West

First host record, nesting behavior, and taxonomic position of the spider wasp genus Hesperopompilus Evans and some other Evans genera (Hymenoptera: Pompilidae), Frank E. Kurczewski, Rick C. West, and James P. Pitts

First host record for the spider wasp Cryptocheilus severini Banks (Hymenoptera: Pompilidae: Pepsinae), Frank E. Kurczewski, Rick C. West, James P. Pitts, and Cecilia Waichert

Additional new and unusual host records for Western Hemisphere spider wasps (Hymenoptera: Pompilidae), Frank E. Kurczewski, Rick C. West, Cecilia Waichert, and James P. Pitts

New records of invasive aphids (Hemiptera: Aphididae) on garlic mustard in the USA, Doris Lagos-Kutz, Rebecah Troutman, and Glen L. Hartman

Synopsis of the subfamily Carventinae in New Zealand (Heteroptera: Aradidae), Marie-Claude Larivière and André Larochelle

Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae), André Larochelle and Marie-Claude Larivière

Whiteflies (Hemiptera: Aleyrodidae) intercepted on plant product imported to South Korea from 2013–2021, Suhyeon Lee and Soo-Jung Suh

Revision of Aneflomorpha Casey and Neaneflus Linsley (Coleoptera: Cerambycidae) of the United States with an illustrated key to species, Steven W. Lingafelter

Three new species of Cautethia Grote (Lepidoptera: Sphingidae) from the Lucayan Archipelago and keys to West Indies species, Jacqueline Y. Miller, Deborah L. Matthews, and Riley J. Gott

Revision of the high Andean genus Eremophygus Ohaus (Coleoptera: Scarabaeidae: Rutelinae: Rutelini), José Mondaca

A new genus for Stromatium chilensis Cerda, 1968 (Coleoptera: Cerambycidae: Cerambycinae: Hesperophanini) from Chile, José Mondaca and Marcos Beéche

First record of Ancognatha erythrodera (Blanchard, 1846) (Coleoptera: Scarabaeidae: Dynastinae: Cyclocephalini) in the altiplano of Chile, José Mondaca and Sergio Rothmann

New fossil false click-beetles from the Americas (Coleoptera: Eucnemidae), Jyrki Muona

Tomoxia bucephala A. Costa (Coleoptera: Mordellidae), a Palearctic tumbling flower beetle established in North America, R. F. C. Naczi, Robert A. Androw, and John Rosenfeld

A new species of Epeolus Latreille, 1802 (Hymenoptera: Apidae) from western North America, Thomas M. Onuferko and Cory S. Sheffield

Revision of the checkered beetle genus Onychotillus Chapin (Coleoptera: Cleridae: Tillinae), Weston Opitz

Megalurothrips usitatus Bagnall (Thysanoptera: Thripidae), first record of an important new pest in Honduras, Jesus Orozco

A new species and new records for two other exotic species of Dirrhagofarsus Fleutiaux, 1935 (Coleoptera: Eucnemidae: Melasinae: Dirhagini) in the United States, Robert L. Otto

New state records for 33 species of Nearctic false click beetles (Coleoptera: Eucnemidae), Robert L. Otto

Culicoides Latreille and Leptoconops Skuse biting midges of the southwestern United States with emphasis on the Canyonlands of southeastern Utah (Diptera: Ceratopogonidae), Robert A. Phillips

Notas sobre Ectinogonia chalyboeiventris chalyboeiventris Germain y Kerremans y E. chalyboeiventris wagenknechti Cobos, con descripción de dos nuevas especies de Ectinogonia Spinola (Coleoptera: Buprestidae) del norte de Chile, Cristian Pineda and José Mondaca

A new species of Neostenoptera Meunier (Diptera: Cecidomyiidae: Winnertziinae) from Hawai‘i, John D. Plakidas, Nhu H. Nguyen, and Michael L. Ferro

New distributional records for Mexican Cleridae (Coleoptera) with the description of three new species, Jacques Rifkind

A new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) from the Sierra Madre Occidental of Mexico, with notes on the type locality of Chrysina adelaida (Hope, 1841), David C. Robacker, David C. Hawks, Naidely Castillo, and Brianna Miranda

A segmented and clawed male foreleg in a newly described genus and species of eumaeine butterfly (Lepidoptera: Lycaenidae), Robert K. Robbins, Christophe Faynel, Jason P.W. Hall, and Robert C. Busby

A checklist of the hawkmoths (Lepidoptera: Sphingidae) of the Cayman Islands: with implications for the pollination of the ghost orchid Dendrophylax fawcettii Rolfe (Orchidaceae: Angraecinae) and consideration of bat predation, M. C. Rose-Smyth, R. R. Askew, P. A. van B. Stafford, M. J. Simon, G. J. Gross, and R. Rozycki

New species, new synonymies, a new rank, and new records in Desmiphorini, Saperdini, and Hemilophini (Coleoptera: Cerambycidae: Lamiinae), Antonio Santos-Silva and Robert A. Androw

Lasioglossum (Dialictus) semicaeruleum (Cockerell, 1895) (Hymenoptera: Halictidae) in Maryland: A disjunct population in eastern North America?, Eugene J. Scarpulla

A new species of Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) from Puerto Rico, Kyle E. Schnepp

Extinct or extant? A new species of Termitodius Wasmann, 1894, (Coleoptera: Scarabaeidae: Aphodiinae: Rhyparini) with a short review of the genus, Paul E. Skelley, Julian Clavijo-Bustos, and Oliver Keller

A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini), Paul E. Skelley and Oliver Keller

In Memoriam: Robert E. Woodruff, July 20, 1933–July 8, 2021, Paul E. Skelley and Oliver Keller

Review of Pharaxonotha Reitter (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting the cycad genus Dioon Lindl. (Cycadales), with descriptions of nine new species and comments on P. kirschii Reitter, Paul E. Skelley, William Tang, and Miguel Angel Pérez-Farrera

Dichotomius woodruffi, a new Dichotomius species of the agenor group from Costa Rica and Nicaragua (Coleoptera: Scarabaeidae: Scarabaeinae), Ángel Solis and Burt Kohlmann

A new genus of predatory midge in the Monohelea complex from Eocene Baltic amber (Diptera: Ceratopogonidae), Ryszard Szadziewski, William L. Grogan Jr., Elżbieta Sontag, and Błażej Bojarski

A new Phyllophaga (Listrochelus>/i>) timida group species from Baja California Sur (Coleoptera: Scarabaeidae: Melolonthinae), William B. Warner

Acmaeodera moesta Dugès, 1891 (Coleoptera: Buprestidae), a poorly known species from Mexico, Richard L. Westcott and Jesus Romero-Napoles

Taxonomic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera), Jing Zhang, Qian Cong, Jinhui Shen, and Nick V. Grishin


Taxonomic revision of Eunota togata (LaFerté-Sénectère, 1841) (Coleoptera: Cicindelidae) in North America with a new subspecies from western Texas and New Mexico, United States, Robert E. Acciavatti

New species, new records and synonymy of Brazilian species of Hypothenemus Westwood, 1834 (Coleoptera: Curculionidae: Scolytinae), Thomas H. Atkinson and Carlos A.H. Flechtmann

The rare beetle egg parasitoid Platystasius transversus (Thomson) (Hymenoptera: Platygastridae: Sceliotrachelinae), a new genus and species record for Germany, Jessica Awad, Cristina Vasilița, and Lars Krogmann

A new Vaejovis C.L. Koch from the Sierra Madre Occidental of Durango, Mexico (Scorpiones: Vaejovidae), Jessica S. Azzinnari, Robert W. Bryson, Matthew R. Graham, Carlos Solís-Rojas, and W. David Sissom

The strange case of Colophotia miranda Olivier, 1886 (Coleoptera: Lampyridae: Luciolinae), Lesley Ballantyne and Wan F.A. Jusoh

Emarginoptyx Ballantyne (Coleoptera: Lampyridae: Luciolinae), a replacement name for the firefly genus Emarginata Ballantyne, 2019, Lesley Ballantyne and Oliver Keller

First record of Palonica viridia (Ball, 1903) (Hemiptera: Membracidae) in Utah, Lawrence Barringer

First record of Phaenops obtusa (Horn) (Coleoptera: Buprestidae) in Massachusetts, Lawrence Barringer

A new genus and species of tiger beetle, Pseudocollyris shooki (Coleoptera: Cicindelidae), from Yunnan, China, Wen-Xuan Bi and Jürgen Wiesner

Una nueva especie de mosca de ojos pedunculados del género Richardia Robineau-Desvoidy y un nuevo reporte de Plagiocephalus latifrons Wiedemann para la costa pacífica de Colombia (Diptera: Tephritoidea: Richardiidae y Ulidiidae), Luis Miguel Constantino

Recharacterization of Stenaspis Audinet-Serville, 1834 with a new species from Mexico (Coleoptera: Cerambycidae: Cerambycinae: Trachyderini), Bryan K. Eya

A new species and significant range extension for the genus Fractipes Bechynĕ (Coleoptera: Chrysomelidae: Eumolpinae: Eumolpini), Wills Flowers

Two new species of Deromecus Solier from Guatemala (Coleoptera: Elateridae: Pomachiliini), E. R. Fuller

Especies nuevas de Lasioglossum Curtis de las altas montañas de las Antillas Mayores y estado actual del conocimiento del taxon Habralictellus Moure y Hurd (Hymenoptera: Apoidea: Halictidae), Julio A. Genaro

New adult host records for three Buprestidae (Coleoptera) rarely encountered in the United States and significant extension of the known geographic range of Agrilus pilosicollis Fisher, Jason A. Hansen, , and Christine A. Nalepa

Checklist of Dermestidae (Insecta: Coleoptera: Bostrichoidea) of the United States, Jiří Háva and Andreas Herrmann

Two new species of palm-leaf skeletonizers (Lepidoptera: Pterolonchidae: Homaledra Busck), James E. Hayden

Two new species of Nothochodaeus Nikolajev, 2005 from Palawan Island, Philippines (Coleoptera: Scarabaeoidea: Ochodaeidae), Jean-Bernard Huchet

A new Neotropical predaceous midge in the genus Parabezzia Malloch from Guerrero, Mexico, and a second Mexican record of P. alexanderi Wirth (Diptera: Ceratopogonidae: Ceratopogoninae: Ceratopogonini), Herón Huerta, Gustavo R. Spinelli, and William L. Grogan Jr.

Survey of the Attelabidae of Wisconsin (Coleoptera: Curculionoidea), Julia Janicki and Daniel K. Young

Luciola lambkinae Keller and Ballantyne, new replacement name for Luciola roseicollis Pic, 1935 (Coleoptera: Lampyridae), Oliver Keller and Lesley Ballantyne

Nocturnal multi-species roosts of Cicindelidae (Coleoptera) in a Neotropical lowland rainforest, Susan Kirmse and Jürgen Wiesner

Taxonomic revision of Cosmodela duponti (Dejean), Cosmodela barmanica (Gestro), new status, and Cosmodela indica (Fleutiaux), new status (Coleoptera: Cicindelidae), Miroslav Klícha and Jürgen Wiesner

Nesting behavior, ecology, and functional morphology of the trapdoor spider-hunting spider wasp Aporus (Plectraporus) hirsutus (Banks) (Hymenoptera: Pompilidae), Frank E. Kurczewski, Alice J. Abela, and Rick C. West

Survivorship of soybean aphid biotypes (Hemiptera: Aphididae) on winter hosts, common and glossy buckthorn, Doris M. Lagos-Kutz and Glen L. Hartman

The lycaenid butterfly fauna (Lepidoptera) of Cosñipata, Peru: annotated checklist, elevational patterns, and rarity, Gerardo Lamas, Michael L. McInnis, Robert C. Busby, and Robert K. Robbins

Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae), André Larochelle and Marie-Claude Larivière

New country records, annotated checklist and key to the dacine fruit flies (Diptera: Tephritidae: Dacinae: Dacini) of Bangladesh, Luc Leblanc, M. Aftab Hossain, Mahfuza Momen, and Kajla Seheli

Occurrence of an exotic whitefly, Siphoninus phillyreae (Haliday) (Hemiptera: Aleyrodidae) in South Korea and its potential pathway analysis, Seon Woo Lee, Hyun Jik Lee, Jun-Ran Kim, and Soo-Jung Suh

A new genus Cicatrisphaerion, new species, new records, and redescriptions of Neotropical Cerambycidae (Coleoptera), Steven W. Lingafelter, Roy F. Morris II, Frederick W. Skillman Jr., and Antonio Santos-Silva

Ishtarella thailandica Martens, new genus, new species (Hymenoptera: Braconidae: Aphidiinae) of aphid parasitoid from Thailand, with a country checklist of Aphidiinae, Abigail P. Martens, Matthew L. Buffington, Donald L.J. Quicke, Marisa Raweearamwong, Buntika A. Butcher, and Paul J. Johnson

How to distinguish the Xestotrachelus Bruner, 1913 (Orthoptera: Romaleidae: Romaleini) from other Romaleini in South America, with a report of the first record in Minas Gerais, Brazil, Daniela Santos Martins Silva, Marcelo Ribeiro Pereira, and Raysa Martins Lima

First record of Orsilochides scurrilis (Stål) (Hemiptera: Heteroptera: Scutelleridae: Pachycorinae) in the United States, with notes on the biology and distribution of U.S. species of Orsilochides Kirkaldy, A. W. Meeds, Joseph E. Eger Jr., and S. D. Carnahan

How do Vanhorniidae (Hymenoptera) parasitize Eucnemidae (Coleoptera)?, Jyrki Muona

The first record of the jumping bristletail Pedetontus gershneri Allen, 1995 (Microcoryphia: Machilidae) from Alabama, USA significantly extends its known range, Alexander B. Orfinger and Grant D. De Jong

A new species of Coomanius Fleutiaux, 1924 (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) from Southeast Asia, Robert L. Otto

A review of the Nearctic species of Nematodes (Coleoptera: Eucnemidae: Macraulacinae: Nematodini) with a description of one new species, Robert L. Otto

Ardella magnaemirabilis (Coleoptera: Scarabaeidae: Melolonthinae: Ardellini), a new scarabaeoid species, genus and tribe from the southwestern United States, M. J. Paulsen

First reports of species-specific ant resemblance in heteronotine treehoppers (Hemiptera: Membracidae: Heteronotinae), Robert Perger