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The holotype of the species, Domninoides mimicus, is based on one tooth. Described in the present report is material containing a practically complete dentition and many postcranial elements which definitely belong with the teeth. This species is closely related to Domninoides valentinensis described by Reed (1962), but some of the antemolars in valentinensis are misidentified. The tooth formulas for valentinensis and for mimicus are given. Development of the cingula on the upper and lower molars in mimicus is conspicuous. Because of similar features in the postcranial bones, the Clarendonian mole may be as fossorial as Scalopus, probably the most fossorial extant North American mole.