Genomic analysis reveals new subspecies of Cercyonis Scudder, 1875 (Nymphalidae: Satyrinae: Satyrini)

Authors

• Jing Zhang, University of Texas Southwestern Medical Center
• Qian Cong, University of Texas Southwestern Medical Center
• Jinhui Shen, University of Texas Southwestern Medical Center
• Leina Song, University of Texas Southwestern Medical Center
• Todd L. Stout, McGuire Center for Lepidoptera and Biodiversity
• Michael S. Fisher
• Robb Hannawacker
• Nick V. Grishin, University of Texas Southwestern Medical CenterFollow

Date of this Version

9-8-2026

Document Type

Journal Issue

Citation

Zhang, J., Q. Cong, J. Shen, L. Song T. L. Stout, M. S. Fisher, R. Hannawacker, & N. V. Grishin.  2026.  Genomic analysis reveals new subspecies of Cercyonis Scudder, 1875 (Nymphalidae: Satyrinae: Satyrini).  The Taxonomic Report 14(2): 1-36. 

Abstract

Genomic analysis of Cercyonis (Cercyonis) Scudder, 1875 (Nymphalidae Rafinesque, 1815) partitions the subgenus into three species groups and reveals five new subspecies from the USA (type localities in parentheses): Cercyonis (Cercyonis) pegala koosharem Stout and Grishin, ssp. n. (Utah, Sevier Co.), Cercyonis (Cercyonis) pegala lasala Grishin, M. Fisher, and Hannawacker ssp. n. (Utah, Grand Co.), Cercyonis (Cercyonis) pegala hortoni Grishin, ssp. n. (Arizona, Gila Co.), Cercyonis (Cercyonis) meadii maculana Grishin, ssp. n. (Colorado, Montezuma Co.), and Cercyonis (Cercyonis) meadii rimana Grishin, ssp. n. (Arizona, Yavapai Co.). A neotype is designated for Satyrus boopis Behr, 1864 (USA, California, Contra Costa Co.). The neotype of Satyrus behrii F. Grinnell, 1905 is invalid and not warranted. A provisional ordered taxonomic list of the subgenus Cercyonis is provided, recognizing six species and 35 subspecies (excluding the nominotypical subspecies).