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The Taxonomic Report of the International Lepidoptera Survey

Lepidoptera Survey

The Taxonomic Report of the International Lepidoptera Survey

The Taxonomic Report (TTR) is the main scientific publication of The International Lepidoptera Survey (TILS). TILS-TTR is currently on hiatus but formerly was published at the rate of 8-10 issues a year. Subscription / dues are usually $45 US for USA and Canada, $55 US for overseas subscribers. At the end of each Volume year, subscribers receive that year's volume on a read-only compact disc for archiving and reproduction for personal use (i.e. a (subscribed) museum or university may make as many copies as needed in whatever format desired). Non-members may receive individual issues in print any time for $10 per issue. Individual issues on CDR to non-members are $25 per issue post paid. Subscriptions and individual issue orders should be made payable to TILS and checks mailed to: Harry Pavulaan, TILS Editor, P.O. Box 1124, Herndon, VA 20172.

2022 2022

Life History and Ecology of the San Emigdio Blue Butterfly (Lepidoptera: Lycaenidae), Gregory R. Ballmer

Table of Contents, Taxonomic Report of the International Lepidoptera Survey, October 1, 1998–December 20, 2022, International Lepidoptera Survey

A New Species of Celastrina from the Northwestern United States and Southwestern Canada with a Lectotype Designation of Lycaena pseudargiolus var. nigrescens Fletcher (Lycaenidae: Polyommatinae), Caitlin C. LaBar, Jonathan P. Pelham, and Norbert G. Kondla

Natural History of the Guatemalan Copper Iophanus pyrrhias (Godman and Salvin, 1887) (Lepidoptera: Lycaenidae) in Guatemala, José Monzón-Sierra and Gregory C. Ballmer

Subspecific Designation for the Central Appalachian Mountains Population of Argynnis (Speyeria) atlantis (W. H. Edwards), Harry Pavulaan

Notes on Eastern North American Lepidoptera, Harry Pavulaan and Annette Allor

Establishment of the Invasive Cactus Moth, Cactoblastis cactorum (Berg) (Lepidoptera: Pyralidae) in Pakistan: A Potential Threat to Cultivated, Ornamental and Wild Opuntia spp. (Cactaceae), Muhammad Ather Rafi, Harry Pavulaan, Muhammad Islam, Muhammad Ashfaq, Haseeb Kamran, Walija Fayaz, Gul Naz Parveen, Riffat Sultana, Ahmad Zia, Waqar Ahmed, Qudrat Ullah, Muhammad Qasim, Falak Naz, Nazeer Ahmed, Muhammad Tariq Khan, Muhammad Saeed, and Jalal Hayat Khan

Genomic DNA Sequencing Reveals Two New North American Species of Staphylus (Hesperiidae: Pyrginae: Carcharodini), Jing Zhang

Neotype Designation for Papilio fulgerator Walch, 1775 (Hesperiidae: Eudaminae), Jing Zhang, Qian Cong, Gerardo Lamas, and Nick V. Grishin

Checking the Checkered Taxonomy of Plötz's Checkered Skippers (Hesperiidae: Pyrgini), Jing Zhang, Qing Cong, John M. Burns, and Nick V. Grishin

Taxonomic Discoveries Enabled by Genomic Analysis of Butterflies, Jing Zhang, Qing Cong, Jinhui Shen, Leina Song, Riley J. Gott, Pierre Boyer, Crispin S. Guppy, Steve Kohler, Gerardo Lamas, Paul A. Opler, and Nick V. Grishin

2021 2021

Two New Species of Hermeuptychia from North America and Three Neotype Designations (Nymphalidae: Satyrinae), Qian Cong, Eduardo P. Barbosa, Mario A. Marín, André V. L. Freitas, Gerardo Lamas, and Nick V. Grishin

Notes on Eastern North American Butterflies, David J. Hoag, Harry Pavulaan, Brian Taber, Kenneth Lorenzen, and Richard D. Ullrich

Review of Montana Euphilotes Mattoni, [1978], with Descriptions of New Taxa (Lepidoptera: Lycaenidae: Polyommatinae), Steve Kohler and Andrew D. Warren

Type Locality of Polyommatus lucia (W. Kirby, 1837) (Lycaenidae: Polyomatinae) with an Inspection of the Species' Phenotypic Expression, Norbert G. Kondla, Harry Pavulaan, and David M. Wright

Evaluation of the Taxonomic Status of Eurytides marcellus form “floridensis” (W. Holland, 1898) (Papilionoidea, Papilioninae, Leptocircini), Harry Pavulaan

Reevaluation of the Described Subspecies of Euphydryas phaeton (Drury, 1773) with a Replacement Name for Melitaea phaeton schausi (Clark, 1927), Harry Pavulaan

Subspecific Designation of the U.S.A. Interior Highlands Population of Argynnis (Speyeria) diana (Cramer, 1777) (Nymphalidae: Heliconiinae: Argynnini: Argynnina), Harry Pavulaan

Egg Plastron of the Bog Copper Butterfly Tharsalea (Epidemia) epixanthe (Bsd. & Le C. [1835]) (Lycaenidae: Lycaeninae), David M. Wright

Pupal Polymorphism of the Bog Copper Butterfly Lycaena (Epidemia) epixanthe (Bsd. & Le C. [1835]) (Lycaenidae: Lycaeninae) in New Jersey, David M. Wright

Genomics-Guided Refinement of Butterfly Taxonomy, Jing Zhang, Qian Cong, Jinhui Shen, Paul A. Opler, and Nick V. Grishin

2020 2020

Review of the Speyeria egleis Complex in Montana, with the Description of Two New Subspecies (Lepidoptera: Nymphalidae: Heliconiinae), Steve Kohler

A Phenotypic Comparison of Regional Populations of Hemileuca maia (Drury, 1773) with Designations of New Subspecies (Bombycoidea, Saturniidae, Hemileucinae), Harry Pavulaan

Designation of Neotype of Hemileuca maia (Drury, 1773) and Refinement of Its Type Locality (Bombycoidea, Saturniidae, Hemileucinae), Harry Pavulaan

Re-evaluation of Euchloe olympia (W. H. Edwards, 1871 (Pieridae: Pierinae) from a Geographic and Taxonomic Perspective, Harry Pavulaan

Genomic Evidence Suggests Further Changes of Butterfly Names, Jing Zhang, Qian Cong, Jinhui Shen, Pal A. Opler, and Nick V. Grishin

2019 2019

Expanded Phenotypic Diagnoses for 24 Recently Named New Taxa of Hesperiidae (Lepidoptera), Nick V. Grishin

Designation of a Neotype for Oeneis melissa (Fabricius, 1775) and a Lectotype for Oeneis polixenes (Fabricius, 1775) (Lepidoptera: Nymphalidae: Satyrinae), Crispin S. Guppy

Changes to North American Butterfly Names, Jing Zhang, Qian Cong, Jinhui Shen, Paul A. Opler, and Nick V. Grishin

2014 2014

Cercyonis pegala agawamensis (Satyridae): A New Butterfly Subspecies from the Coastal Salt Marshes of the Northeastern United States of America, Matthew D. Arey and Alex Grkovich

Larval Host Plants of Enodia anthedon, Satyrodes appalachia and S. eurydice in Vermont, USA, David J. Hoag

A Case of Sympatric Celastrina ladon (Cramer), Celastrina lucia (W. Kirby) and Celastrina neglecta (Edwards) (Lycaenidae: Polyommatinae) in Northern Virginia, with Additional Records of C. lucia in Virginia, Harry Pavulaan

2010 2010

The Nomenclatural Status of Ten Names in the Genus Pieris (Lepidoptera: Pieridae), Crispin S. Guppy and Norbert G. Kondla

Observations on Anthocharis julia browningi and Anthocharis thoosa thoosa Including Tension Zones near Nephi, Juab County, Utah, Todd Stout

Tips on Collecting and Rearing Immatures of 375 Butterfly and Skipper Taxa, Jacque Wolfe, Jack Harry, and Todd Stout

2009 2009

Natural Life Histories of Alaska Colias (Lepidoptera: Pieridae), Jack L. Harry

2007 2007

Yosemite Butterflies (Text), 2nd edition, Kenneth E. Davenport

The Yosemite Butterflies: Color Plates, Kenneth E. Davenport, Norbet G. Kondla, Charles Grisham, and Howard Grisham

A New Subspecies of Colias gigantea from Arctic Alaska (Pieridae), Jack L. Harry

A Description of a New Subspecies of Lycaena phlaeas (Lycaenidae: Lycaeninae) from Montana, United States, With a Comparative Study of Old and New World Populations, Steve Kohler

Bionomic Studies of Papilio demoleus Linnaeus, the Citrus Butterfly, (Lepidoptera: Papilionidae), from Lower Sindh, Pakistan, Arshad Munir, Nikhat Yasmin Siddiqui, Muhammad A. Rafi, Harry Pavulaan, and David Wright

2005 2005

Taxonomic Analysis of the Genus Megisto (Satyridae) in the Eastern United States, Part I: Types, Type Localities, and Taxonomic Relationships of the Available Names, Ronald R. Gatrelle

Immature Stages of Colias johanseni from Arctic Canada (Lepidoptera: Pieridae), Jack Harry

Immature Stages of Colias occidentalis sullivani from Oregon (Lepidoptera: Pieridae), Jack Harry

Immature Stages of Estigmene acrea from Guatemala (Lepidoptera: Arctiidae), Jack Harry

Celastrina serotina (Lycaenidae: Polyommatinae): A New Butterfly Species from the Northeastern United States and Eastern Canada, Harry Pavulaan and David M. Wright

2004 2004

A Concise Update of the Information Provided in The Butterflies of Southern California (1973) by Thomas C. Emmel and John F. Emmel, Kenneth E. Davenport

The Yosemite Butterflies (Text), and The Yosemite Butterflies (Text): Supplement of Additional Information, Collections and Corrections for Volume 5, No. 1, Kenneth E. Davenport

A Taxonomic Examination of Harkenclenus titus (Lycaenidae: Theclinae) in the Eastern United States: Recognition and Resolution of Taxonomic Problems by the Delineation of Old Names, Naming of a Long Recongnized (but Heretofore Undescribed) Subspecies, and Description of a Wide Ranging New Subspecies, Ronald R. Gatrelle

Description of a Multilevel Cryptic New Species of Phyciodes (Nymphalidae: Melitaeinae) from the Southern Appalachian Mountains, Ronald R. Gatrelle

Discovery of a Black Female Form of Pterourus appalachiensis (Papilionidae: Papilioninae) and Additional Observations of the Species in West Virginia, Harry Pavulaan and David M. Wright

2003 2003

A Taxonomic Review of Chlosyne ismeria with Description of a New Subspecies from the Southern Appalachian Mountains, Ronald R. Gatrelle

A Subspecific Assessment of the Genus Hesperia (Hesperiinae) in Eastern North America (Part I: The South): New Subspecies of Hesperia metea, Hesperia sassacus and Hesperia meskei, Ronald R. Gatrelle, Marc C. Minno, and Alex Grkovish

A Case for Taxonomic Recognition of the Taxon Enodia anthedon borealis A. H. Clark (Satyridae), Alex Grkovich and Harry Pavulaan

Fixation of Type Locality for Lycaena acmon Westwood and Characterization of the Species and its Distribution, Paul A. Opler

Butterflies of the North Okanagan, British Columbia, Canada, David L. Threatful

2002 2002

Taxonomy and Distribution of Butterflies (Papilionoidea) of the Skardu Region, Pakistan, Muhammad Abbas, Muhammad Ather Rafi, Mian Inayatullah, Muhammad Rafique Khan, and Harry Pavulaan

A Review of Poanes hobomok (Hesperiidae: Hesperiinae), with the Description of a New Subspecies from the Southern Appalachians, Ronald R. Gatrelle

Name-Bearing Types and Taxonomic Synopsis of Three Lycaenid Butterfly Taxa from Western Canada (Lepidoptera: Lycaenidae), Norbert G. Kondla and Crispin S. Guppy

Pterourus appalachiensis (Papilionidae: Papilioninae), a New Swallowtail Butterfly from the Appalachian Region of the United States, Harry Pavulaan and David M. Wright

2001 2001

An Examination of Southeastern U.S. Satyrium (Lycaenidae: Theclinae), Part One: An Obscure New Subspecies of Satyrium edwardsii, Ronald R. Gatrelle

An Examination of Southeastern U. S. Satyrium (Lycaenidae: Theclinae), Part Two: The Identification and Delimitation of Nominate Satyrium liparops and the Description of a New Subspecies from West Central Peninsular Florida, Ronald R. Gatrelle

An Examination of the Mitoura (Lycaenidae) in the Southeastern United States: With the Description of a New Subspecies of Mitoura hesseli, Ronald R. Gatrelle

Initial Survey of the Butterflies and Skippers in the Vicinity of the Buck Creek Serpentine Barrens, Clay County, North Carolina, Ronald R. Gatrelle

Clarification of and Comments on Northern Speyeria hydaspe Subspecies (Lepidoptera: Nymphalidae), Norbert G. Kondla

2000 2000

A New Species of Angulopis (Lycaenidae, Eumaeini) from Relict Coastal Forest in East-Central Argentina, Gustavo Canals and Kurt Johnson

Records for the Utilization of Prunus as a Larval Foodplant by 71 Species of Lepidoptera in Northeast California, Laurence L. Crabtree and Ron Leuschner

A New North American Swallowtail Butterfly: Description of a Relict Subspecies of Pterourus troilus (Papilionidae) from the Southern Tip of Florida, Ronald R. Gatrelle

Description of a New Subspecies of Poanes aaroni (Hesperioidae: Hesperiinae) from the West Central Gulf Coast of the Southern United States, Ronald R. Gatrelle

A New Elfin Butterfly (Lycaenidae: Eumaeini) from Northern China with Comments on the Nomenclature of Palaearctic Elfins, Kurt Johnson

The Biology, Life History, and Taxonomy of Celastrina neglectamajor (Lycaenidae: Polyommatinae)., Harry Pavulaan and David M. Wright

Nine New Species of Lacinipolia (Noctuidae) from Arizona, California and Vicinity, Charles L. Selman and Ron Leuschner

A Taxonomic Study of, and Key to, the Lecithoceridae (Lepidoptera) from Guizhou, China, Chunsheng Wu

1999 1999

Subspecific Status of Southeastern U.S. Megathymus cofaqui and M. yuccae: Renaming of the Florida Subspecies of M. cofaqui, Ronald R. Gatrele

A Comment on Friedlander’s Asterocampa (Nymphalidae, Apaturinae): Designation of Neotypes for A. celtis and A. clyton, Ronald R. Gatrelle

An Evolutionary Subspecific Assessment of Deciduphagus henrici (Lycaenidae) Based on Its Utilization of Ilex and Non-Ilex Hosts: Description of a Third Ilex Associated Subspecies; Designation of a Neotype and Type Locality for Deciduphagus irus; with Appendix, Ronald R. Gatrelle

Hübner’s helicta, the Forgotten Neonympha: The Recognition and Elevation of Neonympha helicta (Nymphalidae: Satyrinae) to Specific Status, the Designation of Neotypes for N. helicta and N. areolatus, the Subspecific Transfer of septentrionalis to helicta and the Description of a Third helicta Subspecies from South Florida, Ronald R. Gatrelle

Three New Hesperioidae (Hesperiinae) from South Carolina: New Subspecies of Euphyes bimacula, Poanes aaroni, and Hesperia attalus, Ronald R. Gatrelle

A New Subspecies of Brephidium isophthalma (Lycaenidae: Polyommatinae) from Coastal South Carolina, Harry Pavulaan and Ronald R. Gatrelle

Celastrina idella (Lycaenidae: Polyommatinae): A New Butterfly Species from the Atlantic Coastal Plain, David M. Wright and Harry Pavulaan

1998 1998

An Addendum to Anthocharis midea Dos Passos and Klots 1969: Description of a New Subspecies from Texas, Ronald R. Gatrelle

The Rediscovery, Taxonomy, and Biology of Chlosyne gorgone gorgone and Chlosyne ismeria (Nymphalidae) in Burke County, Georgia, Ronald R. Gatrelle

Two New Nymphalidae from Western North Carolina: New Subspecies of Speyeria aphrodite and Phyciodes batesii, Ronald R. Gatrelle