Date of this Version
Published in Wolves: Behavior, Ecology, and Conservation, edited by L. David Mech and Luigi Boitani (Chicago & London: University of Chicago Press, 2003), following page 238.
Adult wolves are very attentive to the pups. Both parents feed and care for them. Any older siblings similarly participate in pup care and feeding. Kin selection is probably the best explanation for the latter behavior. Top: Photograph by Isaac Babcock. Bottom: Photograph by L. David Mech.
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